Tor Carlsen

Section Manager - Botanical Garden
Image of Tor Carlsen
Norwegian version of this page
Phone +47 22851711
Visiting address Sars' gate 1 0562 OSLO
Postal address Postboks 1172 Blindern 0318 OSLO


Cand. Scient. in mycology at the department of Biology, University of Oslo (2002)

PhD in botany at the Botanical garden, University of Oslo (2007)

Academic interests

Fungal- and plant systematics in the genera HelvellaTrichaptumSerpulaSmelowskia, and Cardamine.

Ecology and diversity of root associated fungi, mainly on the plant Bistorta vivipara.


  • Botnen, Synnøve Smebye; Davey, Marie Louise; Aas, Anders Bjørnsgard; Carlsen, Tor; Thoen, Ella; Heegaard, Einar; Vik, Unni; Dresch, Philipp; Mundra, Sunil; Peintner, Ursula; Taylor, Andy F. S. & Kauserud, Håvard (2019). Biogeography of plant root-associated fungal communities in the North Atlantic region mirrors climatic variability. Journal of Biogeography.  ISSN 0305-0270.  46(7), s 1532- 1546 . doi: 10.1111/jbi.13613 Show summary
  • Thoen, Ella; Aas, Anders Bjørnsgard; Vik, Unni; Brysting, Anne Krag; Skrede, Inger; Carlsen, Tor & Kauserud, Håvard (2019). A single ectomycorrhizal plant root system includes a diverse and spatially structured fungal community. Mycorrhiza.  ISSN 0940-6360.  29(3), s 167- 180 . doi: 10.1007/s00572-019-00889-z
  • Cross, Hugh; Sønstebø, Jørn Henrik; Nagy, Nina Elisabeth; Timmermann, Volkmar; Solheim, Halvor; Børja, Isabella; Kauserud, Håvard; Carlsen, Tor; Rzepka, Barbara; Wasak, Katarzyna; Vivian-Smith, Adam & Hietala, Ari Mikko (2017). Fungal diversity and seasonal succession in ash leaves infected by the invasive ascomycete Hymenoscyphus fraxineus. New Phytologist.  ISSN 0028-646X.  213(3), s 1405- 1417 . doi: 10.1111/nph.14204 Full text in Research Archive. Show summary
  • Skrede, Inger; Carlsen, Tor & Schumacher, Trond (2017). A synopsis of the saddle fungi (Helvella: Ascomycota) in Europe – species delimitation, taxonomy and typification. Persoonia.  ISSN 0031-5850.  39, s 201- 253 . doi: 10.3767/persoonia.2017.39.09 Full text in Research Archive.
  • Mysterud, Ivar; Koller, Gry; Høiland, Klaus; Carlsen, Tor & Sletten, Arild (2016). The lamb disease alveld: Search for fungi and bacteria on Narthecium ossifragum foliage and roots. Small Ruminant Research.  ISSN 0921-4488.  136, s 179- 186 . doi: 10.1016/j.smallrumres.2016.01.025
  • Botnen, Synnøve; Kauserud, Håvard; Carlsen, Tor; Blaalid, Rakel & Høiland, Klaus (2015). Mycorrhizal fungal communities in coastal sand dunes and heaths investigated by pyrosequencing analyses. Mycorrhiza.  ISSN 0940-6360.  25(6), s 447- 456 . doi: 10.1007/s00572-014-0624-1
  • Davey, Marie Louise; Blaalid, Rakel; Vik, Unni; Carlsen, Tor; Kauserud, Håvard & Eidesen, Pernille Bronken (2015). Primary succession of Bistorta vivipara (L.) Delabre (Polygonaceae) root-associated fungi mirrors plant succession in two glacial chronosequences. Environmental Microbiology.  ISSN 1462-2912.  17(8), s 2777- 2790 . doi: 10.1111/1462-2920.12770
  • Blaalid, Rakel; Davey, Marie Louise; Kauserud, Håvard; Carlsen, Tor; Halvorsen, Rune; Høiland, Klaus & Eidesen, Pernille Bronken (2014). Arctic root-associated fungal community composition reflects environmental filtering. Molecular Ecology.  ISSN 0962-1083.  23(3), s 649- 659 . doi: 10.1111/mec.12622
  • Botnen, Synnøve; Vik, Unni; Carlsen, Tor; Eidesen, Pernille Bronken; Davey, Marie Louise & Kauserud, Håvard (2014). Low host specificity of root-associated fungi at an Arctic site. Molecular Ecology.  ISSN 0962-1083.  23(4), s 975- 985 . doi: 10.1111/mec.12646
  • Fang, Yao; Vik, Unni; Brysting, Anne Krag; Carlsen, Tor; Halvorsen, Rune & Kauserud, Håvard (2013). Substantial compositional turnover of fungal communities in an alpine ridge-to-snowbed gradient. Molecular Ecology.  ISSN 0962-1083.  22(19), s 5040- 5052 . doi: 10.1111/mec.12437
  • Lindahl, Björn D.; Nilsson, R. Henrik; Tedersoo, Leho; Abarenkov, Kessy; Carlsen, Tor; Kjøller, Rasmus; Kõljalg, Urmas; Pennanen, Taina; Rosendahl, Søren; Stenlid, Jan & Kauserud, Håvard (2013). Fungal community analysis by high-throughput sequencing of amplified markers - a user's guide. New Phytologist.  ISSN 0028-646X.  199(1), s 288- 299 . doi: 10.1111/nph.12243
  • Lindner, Daniel L.; Carlsen, Tor; Nilsson, R. Henrik; Davey, Marie; Schumacher, Trond & Kauserud, Håvard (2013). Employing 454 amplicon pyrosequencing to reveal intragenomic divergence in the internal transcribed spacer rDNA region in fungi. Ecology and Evolution.  ISSN 2045-7758.  3(6), s 1751- 1764 . doi: 10.1002/ece3.586
  • Seierstad, Kristian S.; Carlsen, Tor; Sætre, Glenn-Peter; Hofton, Tom Hellik; Miettinen, Otto & Kauserud, Håvard (2013). A phylogeographic survey of a circumboreal polypore indicates introgression among ecologically differentiated cryptic lineages. Fungal ecology.  ISSN 1754-5048.  6(1), s 119- 128 . doi: 10.1016/j.funeco.2012.09.001
  • Vik, Unni; Logares, Ramiro; Blaalid, Rakel; Halvorsen, Rune; Carlsen, Tor; Bakke, Ingrid; Kolstø, Anne-Brit; Økstad, Ole Andreas Løchen & Kauserud, Håvard (2013). Different bacterial communities in ectomycorrhizae and surrounding soil. Scientific Reports.  ISSN 2045-2322.  3(347) . doi: 10.1038/srep03471 Full text in Research Archive.
  • Blaalid, Rakel; Carlsen, Tor; Kumar, Surendra; Halvorsen, Rune; Ugland, Karl Inne; Fontana, Giovanni & Kauserud, Håvard (2012). Changes in the root-associated fungal communities along a primary succession gradient analysed by 454 pyrosequencing. Molecular Ecology.  ISSN 0962-1083.  21(8), s 1897- 1908 . doi: 10.1111/j.1365-294X.2011.05214.x
  • Carlsen, Tor; Bendiksby, Mika; Hofton, Tom Hellik; Reiso, Sigve; Bakkestuen, Vegar; Haugan, Reidar; Kauserud, Håvard & Timdal, Einar (2012). Species delimitation, bioclimatic range, and conservation status of the threatened lichen Fuscopannaria confusa. The Lichenologist.  ISSN 0024-2829.  44(5), s 565- 575 . doi: 10.1017/S0024282912000199
  • Ikeda, Hajime; Carlsen, Tor; Fujii, Noriyuki; Brochmann, Christian & Setoguchi, Hiroaki (2012). Pleistocene climatic oscillations and the speciation history of an alpine endemic and a widespread arctic-alpine plant. New Phytologist.  ISSN 0028-646X.  194(2), s 583- 594 . doi: 10.1111/j.1469-8137.2012.04061.x
  • Kauserud, Håvard; Kumar, Surendra; Brysting, Anne Krag; Nordén, Jenni & Carlsen, Tor (2012). High consistency between replicate 454 pyrosequencing analyses of ectomycorrhizal plant root samples. Mycorrhiza.  ISSN 0940-6360.  22(4), s 309- 315 . doi: 10.1007/s00572-011-0403-1
  • Skrede, Inger; Carlsen, Tor; Stensrud, Øyvind & Kauserud, Håvard (2012). Genome wide AFLP markers support cryptic species in Coniophora (Boletales). Fungal Genetics and Biology.  ISSN 1087-1845.  116(7), s 778- 784 . doi: 10.1016/j.funbio.2012.04.009
  • Vik, Unni; Carlsen, Tor; Eidesen, Pernille Bronken; Brysting, Anne Krag & Kauserud, Håvard (2012). Microsatellite markers for Bistorta vivipara (Polygonaceae). American Journal of Botany.  ISSN 0002-9122.  99(6), s E226- E229 . doi: 10.3732/ajb.1100504
  • Vik, Unni; Halvorsen, Rune; Ohlson, Mikael; Rydgren, Knut; Carlsen, Tor; Korpelainen, Helena & Kauserud, Håvard (2012). Microsatellite markers for hylocomium splendens (Hylocomiaceae). American Journal of Botany.  ISSN 0002-9122.  99(9), s e344- e346 . doi: 10.3732/ajb.1200028
  • Carlsen, Tor; Engh, Ingeborg Bjorvand; Decock, Cony; Rajchenberg, Mario & Kauserud, Håvard (2011). Multiple cryptic species with divergent substrate affinities in the Serpula himantioides species complex. Fungal Genetics and Biology.  ISSN 1087-1845.  115(1), s 54- 61 . doi: 10.1016/j.funbio.2010.10.004
  • Kumar, Surendra; Carlsen, Tor; Mevik, Bjørn-Helge; Enger, Pål; Blaalid, Rakel; Shalchian-Tabrizi, Kamran & Kauserud, Håvard (2011). CLOTU: An online pipeline for processing and clustering of 454 amplicon reads into OTUs followed by taxonomic annotation. BMC Bioinformatics.  ISSN 1471-2105.  12 . doi: 10.1186/1471-2105-12-182 Full text in Research Archive. Show summary
  • Skrede, Inger; Engh, Ingeborg Bjorvand; Binder, Manfred; Carlsen, Tor; Kauserud, Håvard & Bendiksby, Mika (2011). Evolutionary history of Serpulaceae (Basidiomycota): molecular phylogeny, historical biogeography and evidence for a single transition of nutritional mode. BMC Evolutionary Biology.  ISSN 1471-2148.  11(230) . doi: 10.1186/1471-2148-11-230
  • Bellemain, Eva; Carlsen, Tor; Brochmann, Christian; Coissac, Eric; Taberlet, Pierre & Kauserud, Håvard (2010). ITS as an environmental DNA barcode for fungi: an in silico approach reveals potential PCR biases. BMC Microbiology.  ISSN 1471-2180.  10 . doi: 10.1186/1471-2180-10-189
  • Bjorbækmo, Marit Markussen; Carlsen, Tor; Brysting, Anne Krag; Vrålstad, Trude; Høiland, Klaus; Ugland, Karl Inne; Geml, Jozsef; Schumacher, Trond & Kauserud, Håvard (2010). High diversity of root associated fungi in both alpine and arctic Dryas octopetala. BMC Plant Biology.  ISSN 1471-2229.  10 . doi: 10.1186/1471-2229-10-244
  • Brevik, Ane; Moreno-Garcia, Jaime; Wenelczyk, Joanna; Blaalid, Rakel; Eidesen, Pernille Bronken & Carlsen, Tor (2010). Diversity of fungi associated with Bistorta vivipara (L.) Delarbre root systems along a local chronosequence on Svalbard. Agarica.  ISSN 0800-1820.  29, s 15- 26
  • Carlsen, Tor; Elven, Reidar & Brochmann, Christian (2010). The evolutionary history of Beringian Smelowskia (Brassicaceae) inferred from combined microsatellite and DNA sequence data. Taxon.  ISSN 0040-0262.  59(2), s 427- 438
  • Engh, Ingeborg Bjorvand; Carlsen, Tor; Sætre, Glenn-Peter; Högberg, Nils; Doi, S & Kauserud, Håvard (2010). Two invasive populations of the dry rot fungus Serpula lacrymans show divergent population genetic structures. Molecular Ecology.  ISSN 0962-1083.  19(4), s 706- 715 . doi: 10.1111/j.1365-294X.2009.04505.x
  • Carlsen, Tor; Bleeker, Walter; Hurka, Herbert; Elven, Reidar & Brochmann, Christian (2009). Biogeography and phylogeny of Cardamine (Brassicaceae). Annals of the Missouri Botanical Garden.  ISSN 0026-6493.  96(2), s 215- 236 . doi: 10.3417/2007047
  • Lihova, J; Carlsen, Tor; Brochmann, Christian & Marhold, K (2009). Contrasting phylogeographies inferred for the two alpine sister species Cardamine resedifolia and C-alpina (Brassicaceae). Journal of Biogeography.  ISSN 0305-0270.  36(1), s 104- 120 . doi: 10.1111/j.1365-2699.2008.01998.x
  • Skrede, Inger; Carlsen, Tor; Rieseberg, LH & Brochmann, Christian (2009). Microsatellites for three distantly related genera in the Brassicaceae. Conservation Genetics.  ISSN 1566-0621.  10(3), s 643- 648 . doi: 10.1007/s10592-008-9598-x
  • Jørgensen, Marte Holten; Carlsen, Tor; Skrede, Inger & Elven, Reidar (2008). Microsatellites resolve the taxonomy of the polyploid Cardamine digitata aggregate (Brassicaceae). Taxon.  ISSN 0040-0262.  57(3), s 1- 12
  • Eidesen, Pernille Bronken; Carlsen, Tor; Molau, U & Brochmann, Christian (2007). Repeatedly out of Beringia: Cassiope tetragona embraces the arctic. Journal of Biogeography.  ISSN 0305-0270.  34, s 1559- 1574 . doi: 10.1111/j.1365-2699.2007.01719.x
  • Carlsen, Tor; Sørensen, Steinar & Høiland, Klaus (2004). Sopp på lemen. Blyttia : Norsk botanisk forenings tidsskrift.  ISSN 0006-5269.  62(3), s 183- 185 Show summary
  • Carlsen, Tor & Stensrud, Øyvind (2003). Hattmorkelen Gyromitra longipes Harmaja funnet i Norge. Blyttia : Norsk botanisk forenings tidsskrift.  ISSN 0006-5269.  61(2), s 76- 78 Show summary

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  • Danielsen, Dag Inge; Carlsen, Tor; Finnanger, Anne & Jensen, Ingunn Cecilie (2018). Botanisk hage. Naturhistorisk museum, Universitetet i Oslo.  ISBN 709-00-44-26105-8.  64 s.

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  • Carlsen, Tor (2019). Botanisk hage.
  • Carlsen, Tor (2019, 22. juli). Faretruende arter i norsk flora: Grønnpil og skjørpil. [Radio].  NRK.
  • Carlsen, Tor (2019, 17. juli). Faretruende arter i norsk flora: Høstberberis. [Radio].  NRK.
  • Carlsen, Tor (2019, 15. juli). Faretruende planter i norsk flora: Alaskakornell. [Radio].  NRK.
  • Carlsen, Tor (2019, 18. juli). Faretruende planter i norsk flora: Sprikemispel. [Radio].  NRK.
  • Carlsen, Tor (2019). Hva er en Botanisk hage?.
  • Ørmen, Torfinn; Carlsen, Tor & Gadmar, Tone Charlotte (2018). Hvilke planter og dyr vil fungere i en koloni på Mars? Paneldebatt.
  • Skrede, Inger; Carlsen, Tor & Schumacher, Trond (2015). Artsavgrensning i slekten Helvella ved hjelp av morfologi og molekylære markører ble langt mer utfordrende enn vi hadde regnet med.
  • Skrede, Inger; Carlsen, Tor & Schumacher, Trond (2014). Molecular systematics of the genus Helvella redefines species boundaries and uncover high species diversity.
  • Carlsen, Tor; Aas, Anders Bjørnsgard; Lindner, D; Vrålstad, Trude; Schumacher, Trond & Kauserud, Håvard (2012). Don't make a mista(g)ke: is tag switching an overlooked source of error in amplicon pyrosequencing studies?. Fungal ecology.  ISSN 1754-5048.  5(6), s 747- 749 . doi: 10.1016/j.funeco.2012.06.003 Show summary
  • Tidemann, Grethe; Shalchian-Tabrizi, Kamran; Vrålstad, Trude & Carlsen, Tor (2012, 20. mars). Hvordan skape et toppforskningsmiljø på under fem år.  Uniforum.
  • Nilsson, R. Henrik; Tedersoo, Leho; Lindahl, Björn; Kjøller, Rasmus; Carlsen, Tor; Quince, Christopher; Abarenkov, Kessy; Pennanen, Taina; Stenlid, Jan; Bruns, Tom; Larsson, Karl-Henrik; Kõljalg, Urmas & Kauserud, Håvard (2011). Towards standardization of the description and publication of next-generation sequencing datasets of fungal communities. New Phytologist.  ISSN 0028-646X.  191(2), s 314- 318 . doi: 10.1111/j.1469-8137.2011.03755.x
  • Aas, Anders Bjørnsgard; Carlsen, Tor; Kauserud, Håvard; Mysterud, Atle & Vrålstad, Trude (2010). The effects of animal grazing on species richness and composition of fungal endophytes in a perennial grass. Show summary
  • Aas, Anders Bjørnsgard; Vrålstad, Trude; Carlsen, Tor; Kauserud, Håvard & Mysterud, Atle (2010). Diversity and species composition of fungal endophytes in Avenella flexuosa under different sheep grazing regimes.
  • Carlsen, Tor; Timdal, Einar; Hofton, Tom Hellik; Reiso, Sigve; Mathiesen, Cecilie & Kauserud, Håvard (2009). Phylogenetic species delimitation in Fuscopannaria confusa and implications on its conservation status.
  • Jørgensen, Marte Holten & Carlsen, Tor (2009). Opp av asken, I: Magne Riseng (red.),  Året i bilder 2008.  Cappelen Damm AS.  ISBN 978-82-02-29241-6.  Juni.  s 156 - 161
  • Carlsen, Tor; Kumar, Surendra; Blaalid, Rakel; Brysting, Anne Krag & Kauserud, Håvard (2008). Molecular detection of root-associated fungi Comparing pyrosequencing with Sanger sequencing. Show summary
  • Bjorbækmo, Marit Markussen; Brysting, Anne Krag; Carlsen, Tor; Høiland, Klaus; Geml, Jozsef; Schumacher, Trond; Vrålstad, Trude & Kauserud, Håvard (2008). The ectomycorrhizal fungal community associated with alpine and arctic populations of Dryas octopetala along a latitudinal gradient.
  • Blaalid, Rakel; Carlsen, Tor; Kauserud, Håvard; Brysting, Anne Krag & Høiland, Klaus (2008). Bistorta vivipara and its mycobionts - a model system for ectomycorrhizal research.
  • Lihova, Judith; Carlsen, Tor; Brochmann, Christian & Marhold, Karol (2008). Contrasting phylogeographies inferred for the two alpine sister species Cardamine resedifolia and C. alpina (Brassicaceae).
  • Carlsen, Tor (2007). Are there any trees in the Arctic? Reconstruction of evolutionary histories in a young biome. Series of dissertations submitted to the Faculty of Mathematics and Natural Sciences, University of Oslo.. 665.
  • Carlsen, Tor; Eidesen, Pernille Bronken; Elven, Reidar & Brochmann, Christian (2007). Combining AFLPs, SSRs, and plastid and nuclear sequences in phylogeography and systematics.
  • Carlsen, Tor; Bleeker, Walter; Hurka, Herbert; Elven, Reidar & Brochmann, Christian (2006). Biogeography and phylogeny of Cardamine (Brassicaceae).
  • Carlsen, Tor; Bleeker, Walter; Hurka, Herbert; Albert, Victor Anthony; Elven, Reidar & Brochmann, Christian (2005). Phylogeny and biogeography of Cardamine (Brassicaceae), with emphasis on section Cardaminella.
  • Carlsen, Tor Arne (2003). De norske fylkesblomstene.
  • Carlsen, Tor; Albert, Victor Anthony; Elven, Reidar & Brochmann, Christian (2003). Origin of the modern tundra ecosystem - A new project on comparative phylogenetic analysis of some arctic key genera.
  • Mysterud, Ivar; Høiland, Klaus; Koller, Gry E. B. & Carlsen, Tor Arne (2003). Jakten på alveldens "missing link" (4). Sau og geit.  ISSN 0036-5009.  (3), s 48- 51
  • Mysterud, Ivar; Høiland, Klaus; Koller, Gry; Carlsen, Tor & Sletten, Arild (2003). Jakten på alveldens "missing link" (6). Bakterier på rome i alveldområder i Halsa/Surnadal, Møre og Romsdal 2001. Sau og geit.  ISSN 0036-5009.  56(5), s 40- 41
  • Carlsen, Tor Arne (2002). Fra Linné til DNA forskning. Hvordan arbeider en plantesystematiker i dag?.
  • Carlsen, Tor Arne (2002). Molecular diversity of root endophytes in an alpine Bistorta vivipara - Kobresia myosuroides tundra plant community. Show summary
  • Carlsen, Tor Arne; Høiland, Klaus & Schumacher, Trond (2002). Molecular diversity of root endophytes in a Bistorta vivipara - Kobresia myosuroides tundra plant community. Show summary

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Published Jan. 4, 2018 11:39 AM - Last modified Jan. 10, 2018 2:09 PM